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Homeoboxes

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External resources:

Homeodomain Resource NHGRI

HOX DataBase

Holland PW, Booth HA, Bruford EA. Classification and nomenclature of all human homeobox genes. BMC Biol. 2007 Oct 26;5:47. PMID:17963489

 1. ANTP class homeoboxes
6.    SINE class homeoboxes
HOXL subclass homeoboxes
7.    TALE class homeboxes and pseudogenes
NKL subclass homeoboxes and pseudogenes 8.    CUT class homeoboxes and pseudogenes
2.  PRD class homeoboxes and pseudogenes 9.    PROS class homeoboxes
3.  LIM class homeoboxes 10.  ZF class homeoboxes and pseudogenes
4.  POU class homeoboxes and pseudogenes 11.  CERS class homeoboxes
5.  HNF class homeoboxes  

Click on the Approved Gene Symbol to retrieve the complete gene record

ANTP class homeoboxes: HOXL subclass homeoboxes

Approved Symbol Approved Name Previous Symbols Synonyms Chromosome
HOXA1 homeobox A1 HOX1F, HOX1 7p15.2
HOXA2 homeobox A2 HOX1K 7p15.2
HOXA3 homeobox A3 HOX1E, HOX1 7p15.2
HOXA4 homeobox A4 HOX1D, HOX1 7p15.2
HOXA5 homeobox A5 HOX1C, HOX1 7p15.2
HOXA6 homeobox A6 HOX1B, HOX1 7p15.2
HOXA7 homeobox A7 HOX1A, HOX1 7p15.2
HOXA9 homeobox A9 HOX1G, HOX1 7p15.2
HOXA10 homeobox A10 HOX1H, HOX1 7p15.2
HOXA11 homeobox A11 HOX1I, HOX1 7p15.2
HOXA13 homeobox A13 HOX1J, HOX1 7p15.2
HOXB1 homeobox B1 HOX2, HOX2I 17q21.32
HOXB2 homeobox B2 HOX2, HOX2H 17q21.32
HOXB3 homeobox B3 HOX2, HOX2G 17q21.32
HOXB4 homeobox B4 HOX2, HOX2F 17q21.32
HOXB5 homeobox B5 HOX2, HOX2A 17q21.32
HOXB6 homeobox B6 HOX2, HOX2B 17q21.32
HOXB7 homeobox B7 HOX2, HOX2C 17q21.32
HOXB8 homeobox B8 HOX2, HOX2D 17q21.32
HOXB9 homeobox B9 HOX2E, HOX2 17q21.32
HOXB13 homeobox B13 17q21.32
HOXC4 homeobox C4 HOX3, HOX3E 12q13.13
HOXC5 homeobox C5 HOX3D, HOX3 12q13.13
HOXC6 homeobox C6 HOX3C, HOX3 12q13.13
HOXC8 homeobox C8 HOX3, HOX3A 12q13.13
HOXC9 homeobox C9 HOX3, HOX3B 12q13.13
HOXC10 homeobox C10 HOX3I 12q13.13
HOXC11 homeobox C11 HOX3H 12q13.13
HOXC12 homeobox C12 HOX3, HOX3F, HOC3F 12q13.13
HOXC13 homeobox C13 HOX3, HOX3G 12q13.13
HOXD1 homeobox D1 HOX4, HOX4G 2q31.1
HOXD3 homeobox D3 HOX4A, HOX4, HOX1D 2q31.1
HOXD4 homeobox D4 HOX4B, HOX4 2q31.1
HOXD8 homeobox D8 HOX4, HOX4E 2q31.1
HOXD9 homeobox D9 HOX4C, HOX4 2q31.1
HOXD10 homeobox D10 HOX4, HOX4D 2q31.1
HOXD11 homeobox D11 HOX4, HOX4F 2q31.1
HOXD12 homeobox D12 HOX4H 2q31.1
HOXD13 homeobox D13 HOX4I, SPD 2q31.1
CDX1 caudal type homeobox 1 5q32
CDX2 caudal type homeobox 2 CDX3 13q12.2
CDX4 caudal type homeobox 4 Xq13.2
EVX1 even-skipped homeobox 1 7p15.2
EVX2 even-skipped homeobox 2 2q31.1
GBX1 gastrulation brain homeobox 1 7q36.1
GBX2 gastrulation brain homeobox 2 2q37.2
GSX1 GS homeobox 1 GSH1 Gsh-1 13q12.2
GSX2 GS homeobox 2 Gsh2 4q12
MEOX1 mesenchyme homeobox 1 MOX1 17q21.31
MEOX2 mesenchyme homeobox 2 GAX MOX2 7p22.1-p21.3
MNX1 motor neuron and pancreas homeobox 1 HLXB9 HB9, HOXHB9, SCRA1 7q36
PDX1 pancreatic and duodenal homeobox 1 IPF1 IDX-1, STF-1, PDX-1, MODY4 13q12.1

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ANTP class homeoboxes:  NKL subclass homeoboxes and pseudogenes  

Approved Symbol Approved Name Previous Symbols Synonyms Chromosome
NKX1-1 NK1 homeobox 1 HSPX153, SAX2 4p16.3
NKX1-2 NK1 homeobox 2 C10orf121 bB238F13.2 10q26.13
NKX2-1 NK2 homeobox 1 NKX2A, BCH, TITF1 TTF-1, TTF1 14q13.3
NKX2-2 NK2 homeobox 2 NKX2B NKX2.2 20p11.22
NKX2-3 NK2 homeobox 3 NKX2C NKX2.3, CSX3, NKX4-3 10q24.2
NKX2-4 NK2 homeobox 4 NKX2D NKX2.4 20p11.22
NKX2-5 NK2 homeobox 5 CSX, NKX2E CSX1, NKX2.5, NKX4-1 5q34
NKX2-6 NK2 homeobox 6 CSX2, NKX4-2 8p21.2
NKX2-8 NK2 homeobox 8 NKX2H NKX2.8, Nkx2-9 14q13.3
NKX3-1 NK3 homeobox 1 NKX3A NKX3.1, BAPX2 8p21.2
NKX3-2 NK3 homeobox 2 BAPX1 NKX3B, NKX3.2 4p16.3
NKX6-1 NK6 homeobox 1 NKX6A Nkx6.1 4q21.33
NKX6-2 NK6 homeobox 2 NKX6B, GTX, NKX6.1 10q26.3
NKX6-3 NK6 homeobox 3 FLJ25169 8p11.21
BARHL1 BarH-like homeobox 1 9q34.13
BARHL2 BarH-like homeobox 2 1p22.2
BARX1 BARX homeobox 1 9q22.32
BARX2 BARX homeobox 2 11q24.3
BSX brain-specific homeobox BSX1 11q24.1
DBX1 developing brain homeobox 1 11p15.1
DBX2 developing brain homeobox 2 FLJ16139 12q12
DLX1 distal-less homeobox 1 2q31.1
DLX2 distal-less homeobox 2 TES-1 2q31.1
DLX3 distal-less homeobox 3 17q21.33
DLX4 distal-less homeobox 4 DLX7, DLX9 DLX8, BP1 17q21.33
DLX5 distal-less homeobox 5 7q21.3
DLX6 distal-less homeobox 6 7q21.3
EMX1 empty spiracles homeobox 1 2p13.2
EMX2 empty spiracles homeobox 2 10q26.11
EN1 engrailed homeobox 1 2q14.2
EN2 engrailed homeobox 2 7q36.2
HHEX hematopoietically expressed homeobox PRHX HEX, HOX11L-PEN 10q23.33
HLX H2.0-like homeobox HLX1 HB24 1q41
HMX1 H6 family homeobox 1 H6, NKX5-3 4p16.1
HMX2 H6 family homeobox 2 NKX5-2 10q26.13
HMX3 H6 family homeobox 3 NKX5-1 10q26.13
LBX1 ladybird homeobox 1 LBX1H, HPX6 10q24.32
LBX2 ladybird homeobox 2 2p13.1
MSX1 msh homeobox 1 HOX7 HYD1, OFC5 4p16.2
MSX2 msh homeobox 2 PFM1 CRS2, FPP, HOX8, MSH, PFM 5q35.2
MSX2P1 msh homeobox 2 pseudogene 1 MSX2P HSHPX5, HPX5 17q23.2
NANOG Nanog homeobox FLJ12581, FLJ40451 12p13.31
NANOGP1 Nanog homeobox pseudogene 1 NANOG2 12p13.31
NANOGP2 Nanog homeobox pseudogene 2 NANOGP4 2q36.1
NANOGP3 Nanog homeobox pseudogene 3 6p12.1
NANOGP4 Nanog homeobox pseudogene 4 NANOGP2 7p15.3
NANOGP5 Nanog homeobox pseudogene 5 9q31.1
NANOGP6 Nanog homeobox pseudogene 6 10q24.2
NANOGP7 Nanog homeobox pseudogene 7 NANOGP3 14q32.12
NANOGP8 Nanog homeobox pseudogene 8 15q13.3
NANOGP9 Nanog homeobox pseudogene 9 Xq12
NANOGP10 Nanog homeobox pseudogene 10 Xp11.4
NANOGP11 Nanog homeobox pseudogene 11 6q25
NOTO notochord homeobox 2p13.2
TLX1 T-cell leukemia homeobox 1 TCL3, HOX11 10q24.32
TLX2 T-cell leukemia homeobox 2 HOX11L1 Enx, Tlx2, NCX 2p13.1
TLX3 T-cell leukemia homeobox 3 HOX11L2 RNX 5q35.1
VAX1 ventral anterior homeobox 1 10q26.11
VAX2 ventral anterior homeobox 2 DRES93 2p13.3
VENTX VENT homeobox VENTX2 HPX42B 10q26.3
VENTXP1 VENT homeobox pseudogene 1 NA88A, CT18 Xp21.3
VENTXP2 VENT homeobox pseudogene 2 13q31.1
VENTXP3 VENT homeobox pseudogene 3 12q21.1
VENTXP4 VENT homeobox pseudogene 4 3p24.2
VENTXP5 VENT homeobox pseudogene 5 8p12
VENTXP6 VENT homeobox pseudogene 6 8q21.11
VENTXP7 VENT homeobox pseudogene 7 VENTX1 HPX42 3p24.3

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 PRD class homeoboxes and pseudogenes:

Approved Symbol Approved Name Previous Symbols Synonyms Chromosome
ALX1 ALX homeobox 1 CART1 12q21.31
ALX3 ALX homeobox 3 FND 1p13.3
ALX4 ALX homeobox 4 PFM2 FPP, PFM, KIAA1788 11p11.2
ARGFX arginine-fifty homeobox 3q13.33
ARGFXP1 arginine-fifty homeobox pseudogene 1 5q23.2
ARGFXP2 arginine-fifty homeobox pseudogene 2 17q11.2
ARX aristaless related homeobox MRXS1, PRTS, MRX76, MRX54, MRX43, MRX36, MRX29, MRX32, MRX33, MRX38, MRX87 ISSX, CT121, EIEE1 Xp21.3
CRX cone-rod homeobox CORD2 CRD, LCA7, OTX3 19q13.3
DMBX1 diencephalon/mesencephalon homeobox 1 OTX3 PAXB 1p34.1
DPRX divergent-paired related homeobox 19q13.42
DPRXP1 divergent-paired related homeobox pseudogene 1 2q32.1
DPRXP2 divergent-paired related homeobox pseudogene 2 6p21.31
DPRXP3 divergent-paired related homeobox pseudogene 3 14q13.2
DPRXP4 divergent-paired related homeobox pseudogene 4 17q11.2
DPRXP5 divergent-paired related homeobox pseudogene 5 21q22.13
DPRXP6 divergent-paired related homeobox pseudogene 6 Xp11.4
DPRXP7 divergent-paired related homeobox pseudogene 7 Xq23
DRGX dorsal root ganglia homeobox PRRXL1 DRG11 10q11.23
DUX4L1 double homeobox 4 like 1 DUX4 4q35.2
DUX4L2 double homeobox 4 like 2 4q35.2
DUX4L3 double homeobox 4 like 3 4q35.2
DUX4L4 double homeobox 4 like 4 4q35.2
DUX4L5 double homeobox 4 like 5 4q35.2
DUX4L6 double homeobox 4 like 6 4q35.2
DUX4L7 double homeobox 4 like 7 4q35.2
DUX4L8 double homeobox 4 like 8 DUX2 4q35.2
DUX4L9 double homeobox 4 like 9 4q35.2
DUX4L10 double homeobox 4 like 10 DUX10 10q26.3
DUX4L11 double homeobox 4 like 11 10q26.3
DUX4L12 double homeobox 4 like 12 10q26.3
DUX4L13 double homeobox 4 like 13 10q26.3
DUX4L14 double homeobox 4 like 14 10q26.3
DUX4L15 double homeobox 4 like 15 10q26.3
DUX4L16 double homeobox 4 like 16 DUXY1 Yq11.21
DUX4L17 double homeobox 4 like 17 DUXY3 Yq11.21
DUX4L18 double homeobox 4 like 18 DUXY2 Yq11.21
DUX4L19 double homeobox 4 like 19 DUXY4 Yq11.21
DUXA double homeobox A 19q13.43
DUXAP1 double homeobox A pseudogene 1 2p11.2
DUXAP2 double homeobox A pseudogene 2 8q22.3
DUXAP3 double homeobox A pseudogene 3 10q11.21
DUXAP4 double homeobox A pseudogene 4 10q11.21
DUXAP5 double homeobox A pseudogene 5 11q23.3
DUXAP6 double homeobox A pseudogene 6 15q26.1
DUXAP7 double homeobox A pseudogene 7 20p11.23
DUXAP8 double homeobox A pseudogene 8 22q11.21
DUXAP9 double homeobox A pseudogene 9 14q11.2
DUXAP10 double homeobox A pseudogene 10 14q11.2
DUXB double homeobox B 16q23.1
ESX1 ESX homeobox 1 ESX1L ESXR1 Xq22.2
GSC goosecoid homeobox 14q32.13
GSC2 goosecoid homeobox 2 GSCL 22q11.21
HESX1 HESX homeobox 1 RPX, ANF 3p14.3
HOPX HOP homeobox LAGY, HOP, OB1, NECC1, SMAP31 4q12
ISX intestine-specific homeobox RAXLX 22q12.3
LEUTX leucine twenty homeobox 19q13.2
MIXL1 Mix paired-like homeobox MILD1, MIXL 1q42.12
NOBOX NOBOX oogenesis homeobox OG2, Og2x 7q35
OTP orthopedia homeobox 5q14.1
OTX1 orthodenticle homeobox 1 2p15
OTX2 orthodenticle homeobox 2 14q22.3
OTX2P1 orthodenticle homeobox 2 pseudogene 1 OTX2P 9q21
PAX2 paired box 2 10q24.31
PAX3 paired box 3 WS1 HUP2 2q36.1
PAX4 paired box 4 MODY9 7q32.1
PAX5 paired box 5 BSAP 9p13.2
PAX6 paired box 6 AN2 D11S812E, AN, WAGR 11p13
PAX7 paired box 7 Hup1 1p36.13
PAX8 paired box 8 2q13
PHOX2A paired-like homeobox 2a ARIX, FEOM2 PMX2A, CFEOM2 11q13.4
PHOX2B paired-like homeobox 2b PMX2B Phox2b, NBPhox 4p13
PITX1 paired-like homeodomain 1 BFT PTX1, POTX 5q31.1
PITX2 paired-like homeodomain 2 IRID2, IHG2, RIEG, RIEG1, RGS IGDS, RS, Brx1, Otlx2, ARP1 4q25
PITX3 paired-like homeodomain 3 ASMD 10q24.32
PROP1 PROP paired-like homeobox 1 5q35.3
PRRX1 paired related homeobox 1 PMX1 PHOX1 1q24.3
PRRX2 paired related homeobox 2 PRX2, PMX2 9q34.11
RAX retina and anterior neural fold homeobox RX 18q21.31
RAX2 retina and anterior neural fold homeobox 2 RAXL1 MGC15631, ARMD6, CORD11 19p13.3
RHOXF1 Rhox homeobox family, member 1 OTEX, PEPP1 Xq24
RHOXF2 Rhox homeobox family, member 2 THG1, PEPP-2, PEPP2, CT107 Xq24
RHOXF2B Rhox homeobox family, member 2B Xq24
SEBOX SEBOX homeobox OG9, OG9X 17q11.2
SHOX short stature homeobox PHOG, GCFX, SS, SHOXY Xp22.33 and Yp11.32
SHOX2 short stature homeobox 2 SHOT, OG12X, OG12 3q25.32
TPRX1 tetra-peptide repeat homeobox 1 FLJ40321 19q13.33
TPRX1P1 tetra-peptide repeat homeobox 1 pseudogene 1 10q22.3
TPRX2P tetra-peptide repeat homeobox 2 pseudogene TPRX2P1 19q13.32
TPRXL tetra-peptide repeat homeobox-like FLJ35107 3p25.1
UNCX UNC homeobox Uncx4.1 7p22.3
VSX1 visual system homeobox 1 PPCD PPD, PPCD1 20p11.21
VSX2 visual system homeobox 2 HOX10, CHX10 RET1 14q24.3

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LIM class homeoboxes:

Approved Symbol Approved Name Previous Symbols Synonyms Chromosome
LHX1 LIM homeobox 1 LIM-1, LIM1 17q12
LHX2 LIM homeobox 2 LH-2, hLhx2 9q33.3
LHX3 LIM homeobox 3 9q34.3
LHX4 LIM homeobox 4 Gsh4 1q25.3
LHX5 LIM homeobox 5 12q24.13
LHX6 LIM homeobox 6 LHX6.1 9q33.2
LHX8 LIM homeobox 8 Lhx7 1p31.1
LHX9 LIM homeobox 9 1q31.1
LMX1A LIM homeobox transcription factor 1, alpha LMX1 LMX1.1 1q24.1
LMX1B LIM homeobox transcription factor 1, beta NPS1 9q33.3
ISL1 ISL LIM homeobox 1 Isl-1, ISLET1 5q11.2
ISL2 ISL LIM homeobox 2 FLJ10160 15q24.3

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POU class homeoboxes and pseudogenes:

Approved Symbol Approved Name Previous Symbols Synonyms Chromosome
POU1F1 POU class 1 homeobox 1 PIT1 GHF-1, POU1F1a 3p11.2
POU2F1 POU class 2 homeobox 1 OTF1 OCT1 1q24.2
POU2F2 POU class 2 homeobox 2 OTF2 OCT2 19q13.2
POU2F3 POU class 2 homeobox 3 OCT11, PLA-1, Skn-1a, Epoc-1 11q23.3
POU3F1 POU class 3 homeobox 1 OTF6 OCT6, SCIP 1p34.3
POU3F2 POU class 3 homeobox 2 OTF7 POUF3, BRN2, OCT7 6q16.2
POU3F3 POU class 3 homeobox 3 BRN1, OTF8 2q12.1
POU3F4 POU class 3 homeobox 4 DFN3 BRN4, OTF9, DFNX2 Xq21.1
POU4F1 POU class 4 homeobox 1 BRN3A RDC-1 13q31.1
POU4F2 POU class 4 homeobox 2 BRN3B Brn-3b 4q31.22
POU4F3 POU class 4 homeobox 3 DFNA15 BRN3C 5q32
POU5F1 POU class 5 homeobox 1 OTF3 OCT3, Oct4, MGC22487 6p21.33
POU5F1B POU class 5 homeobox 1B OTF3P1, POU5F1P1 OTF3C 8q24.21
POU5F1P2 POU class 5 homeobox 1 pseudogene 2 8q22.3
POU5F1P3 POU class 5 homeobox 1 pseudogene 3 OTF3L, POU5F1L POU5FLC12 12p13.31
POU5F1P4 POU class 5 homeobox 1 pseudogene 4 POU5FLC1, Oct4-pg4 1q22
POU5F1P5 POU class 5 homeobox 1 pseudogene 5 Oct4-pg5 10q21.3
POU5F1P6 POU class 5 homeobox 1 pseudogene 6 3q21.3
POU5F1P7 POU class 5 homeobox 1 pseudogene 7 3q12.1
POU5F1P8 POU class 5 homeobox 1 pseudogene 8 17q25.3
POU5F2 POU domain class 5, transcription factor 2 SPRM-1, FLJ25680 5q15
POU6F1 POU class 6 homeobox 1 BRN5, MPOU, TCFB1 12q13.13
POU6F2 POU class 6 homeobox 2 RPF-1 7p14.1
HDX highly divergent homeobox CXorf43 FLJ30678 Xq21.1

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 HNF class homeoboxes:

Approved Symbol Approved Name Previous Symbols Synonyms Chromosome
HNF1A HNF1 homeobox A MODY3, TCF1 HNF1, LFB1 12q24.31
HNF1B HNF1 homeobox B TCF2 LFB3, VHNF1, HNF1beta, MODY5 17q12
HMBOX1 homeobox containing 1 HNF1LA, PBHNF, FLJ21616, HOT1 8p12

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 SINE class homeoboxes:

Approved Symbol Approved Name Previous Symbols Synonyms Chromosome
SIX1 SIX homeobox 1 DFNA23 14q23.1
SIX2 SIX homeobox 2 2p21
SIX3 SIX homeobox 3 HPE2 2p21
SIX4 SIX homeobox 4 AREC3 14q23.1
SIX5 SIX homeobox 5 DMAHP 19q13.32
SIX6 SIX homeobox 6 OPTX2 Six9 14q23.1

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 TALE class homeboxes and pseudogenes:

Approved Symbol Approved Name Previous Symbols Synonyms Chromosome
IRX1 iroquois homeobox 1 IRX-5 5p15.33
IRX1P1 iroquois homeobox 1 pseudogene 1 13q12.12
IRX2 iroquois homeobox 2 5p15.33
IRX3 iroquois homeobox 3 IRX-1 16q12.2
IRX4 iroquois homeobox 4 5p15.33
IRX4P1 iroquois homeobox 4 pseudogene 1 18p11.22
IRX5 iroquois homeobox 5 IRX-2a 16q12.2
IRX6 iroquois homeobox 6 IRX7 IRX-3 16q12.2
MEIS1 Meis homeobox 1 2p14
MEIS2 Meis homeobox 2 MRG1, HsT18361 15q14
MEIS3 Meis homeobox 3 MRG2, DKFZp547H236 19q13.32
MEIS3P1 Meis homeobox 3 pseudogene 1 MEIS3, MEIS4 MRG2 17p12
MEIS3P2 Meis homeobox 3 pseudogene 2 17p11.2
MKX mohawk homeobox C10orf48, IRXL1 MGC39616 10p12.1
PBX1 pre-B-cell leukemia homeobox 1 1q23.3
PBX2 pre-B-cell leukemia homeobox 2 G17, HOX12, PBX2MHC 6p21.32
PBX2P1 pre-B-cell leukemia homeobox 2 pseudogene 1 PBX2, PBXP1 3q24
PBX3 pre-B-cell leukemia homeobox 3 9q33.3
PBX4 pre-B-cell leukemia homeobox 4 19p12
PKNOX1 PBX/knotted 1 homeobox 1 PREP1 21q22.3
PKNOX2 PBX/knotted 1 homeobox 2 11q24.2
TGIF1 TGFB-induced factor homeobox 1 HPE4, TGIF 18p11.31
TGIF1P1 TGFB-induced factor homeobox 1 pseudogene 1 19q13.32
TGIF2 TGFB-induced factor homeobox 2 20q11.23
TGIF2LX TGFB-induced factor homeobox 2-like, X-linked Xq21.31
TGIF2LY TGFB-induced factor homeobox 2-like, Y-linked Yp11.2
TGIF2P1 TGFB-induced factor homeobox 2 pseudogene 1 1q44
TGIF2P2 TGFB-induced factor homeobox 2 pseudogene 2 15q21.1
TGIF2P3 TGFB-induced factor homeobox 2 pseudogene 3 15q21.1
TGIF2P4 TGFB-induced factor homeobox 2 pseudogene 4 14q24.2

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 CUT class homeoboxes and pseudogenes:

Approved Symbol Approved Name Previous Symbols Synonyms Chromosome
ONECUT1 one cut homeobox 1 HNF6, HNF6A HNF-6 15q21.3
ONECUT2 one cut homeobox 2 OC-2 18q21.31
ONECUT3 one cut homeobox 3 19p13.3
CUX1 cut-like homeobox 1 CUTL1 CDP, CDP1, CUX, CUT, Clox, CDP/Cut, CDP/Cux, Cux/CDP, CASP, GOLIM6 7q22.1
CUX2 cut-like homeobox 2 CUTL2 KIAA0293, CDP2 12q24.12
CUX2P1 cut-like homeobox 2 pseudogene 1 10p14
CUX2P2 cut-like homeobox 2 pseudogene 2 4q32.1
SATB1 SATB homeobox 1 3p24.3
SATB1P1 SATB homeobox 1 pseudogene 1 6q22.31
SATB2 SATB homeobox 2 KIAA1034, FLJ21474 2q33.1

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 PROS class homeoboxes:

Approved Symbol Approved Name Previous Symbols Synonyms Chromosome
PROX1 prospero homeobox 1 1q41
PROX2 prospero homeobox 2 FLJ36749 14q24.3

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 ZF class homeoboxes and pseudogenes:

Approved Symbol Approved Name Previous Symbols Synonyms Chromosome
ZFHX2 zinc finger homeobox 2 ZNF409 KIAA1762, KIAA1056, ZFH-5 14q11.2
ZFHX3 zinc finger homeobox 3 ATBF1 ZNF927 16q22.3
ZFHX4 zinc finger homeobox 4 ZFH4, FLJ20980 8q21.11
ZHX1 zinc fingers and homeoboxes 1 8q24.13
ZHX2 zinc fingers and homeoboxes 2 KIAA0854 8q24.13
ZHX3 zinc fingers and homeoboxes 3 TIX1 KIAA0395 20q12
ZEB1 zinc finger E-box binding homeobox 1 TCF8, PPCD3 BZP, ZEB, AREB6, NIL-2-A, Zfhep, Zfhx1a, FECD6 10p11.22
ZEB2 zinc finger E-box binding homeobox 2 ZFHX1B KIAA0569, SIP-1, SIP1 2q22.3
ZEB2P1 zinc finger E-box binding homeobox 2 pseudogene 1 ZEB1P1 4p15.32
ADNP activity-dependent neuroprotector homeobox KIAA0784, ADNP1 20q13.13
HOMEZ homeobox and leucine zipper encoding KIAA1443 14q11.2

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CERS class homeoboxes:

Approved Symbol Approved Name Previous Symbols Synonyms Chromosome
CERS2 ceramide synthase 2 LASS2 SP260, FLJ10243 1q21.3
CERS3 ceramide synthase 3 LASS3 MGC27091 15q26.3
CERS4 ceramide synthase 4 LASS4 FLJ12089, Trh1 19p13.3
CERS5 ceramide synthase 5 LASS5 Trh4, MGC45411, FLJ25304 12q13.12
CERS6 ceramide synthase 6 LASS6 2q31

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